**Broad Sense Heritability:** estimates percentage of genetic variance using all components of genetic variance

: ratio of genotypic variance to phenotypic variance

**Narrow Sense Heritability:** estimates percentage of genetic variance using only additive variance

: ratio of additive genotypic variance to phenotypic variance

**Variance Components of a Quantitative Trait:**

**V _{P} = V_{G} + V_{E} + V**

where: **V**_{P} = phenotypic variance**V**_{G} = genetic variation**V**_{E} = environmental variation**V**_{GE} = variation associated with the genetic and environmental factor interactions

Suppose that phenotypic variation in the tail length of mice is summarized in the table below.

What is the narrow-sense heritability (h^{2}) of tail length?

What is the broad-sense heritability (H^{2}) of tail length?

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